Like mycorrhiza, I'd like to see the original Wilf and Ewens paper to know exactly what is being criticised: only the
abstract is available online. I'd also echo his point about Ewert and co. criticising the metaphor rather than the process.
Ewert and co. make great play of biological factors they claim Wilf and Ewens ignore (epistasis, pleiotropy, etc) without once demonstrating that incorporating these factors into the model would make the difference they wish to see - i.e. making the evolutionary process impossibly slow. On the subject of omitted biology, I note Ewert & co make no mention of homeotic mutations, but are hung up on changes in protein-encoding genes. (Nor do Wilf and Ewens refer to homeobox genes in their abstract, but changes in such genes would have a powerful effect on the rate of evolution.)
Indeed; and neither do Ewert & co show that their claimed omissions would make that time impossibly long. What they do is show a nice line in conflation:
The bolding is mine: you will note that the authors are presenting a human-specific calculation as though its outcome applies to all organisms.
The essence of Ewert's argument seems to be that the mathematical model used by Wilf and Ewens is over-simplified. (Perhaps it would be better to say more simplified than it might be, since any mathematical model is by definition a simplification of nature.) I am not enough of a mathematician to judge the rightness of this claim, but would point out that even if (a substantial if) the criticisms are justified, what they have achieved is showing a model supporting the sufficient-time idea to be sub-optimal. Despite the optimistic title given to the thread, that is not the same as demonstrating insufficient time. (If Alice publishes arguments for Shakespeare really being the author of Hamlet, and Bob points out flaws in her arguments, Bob has not thereby demonstrated that someone else wrote the play.)